I have spent thirty years studying the population ecology of red and grey squirrels in Europe and western Canada. My work is currently focussed on pine marten translocation in North Wales.

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

I frequently supervise both undergraduate and postgraduate research, offering local field based studies on Anglesey or in Gwynedd.

My research encompasses viral epidemiological studies with a focus upon Squirrel pox, Adenovirus and Rota virus in sciurid populations; genetic surveillance of red and grey squirrel populations in Europe and North America, wildlife translocation and invasive species management.

I am currently managing the reintroduction of pine marten into North Wales and red squirrel in Clocaenog forest in the North East.

2024

2023

2022

2021

2020

2019

  • PublishedAssessing a potential non-invasive method for viral diagnostic purposes in European squirrels
    Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50
    Research output: Contribution to journal › Article › peer-review
  • PublishedBritish red squirrels remain the only known wild rodent host for leprosy bacilli
    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8
    Research output: Contribution to journal › Article › peer-review
  • PublishedBy trying to be everything, rewilding risks meaning nothing
    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438
    Research output: Contribution to journal › Letter
  • PublishedControlling grey squirrels in forests and woodlands in the UK
    Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.
    Research output: Book/Report › Book › peer-review
  • PublishedDeconstructing compassionate conservation
    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768
    Research output: Contribution to journal › Article › peer-review
  • PublishedDetecting viral infection in red squirrels
    Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Apr 2019, In: Veterinary Record. 184, p. 507
    Research output: Contribution to journal › Article
  • PublishedDisease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23
    Research output: Contribution to journal › Article › peer-review
  • PublishedFirst report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3
    Research output: Contribution to journal › Article › peer-review
  • PublishedLeprosy in red squirrels in the UK
    Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416
    Research output: Contribution to journal › Article
  • PublishedPine marten scat holds few clues for squirrel disease
    Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698
    Research output: Contribution to journal › Article
  • PublishedRed squirrel bark-stripping of hornbeam in North Wales
    Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277
    Research output: Contribution to journal › Article › peer-review
  • PublishedReintroducing rewilding to restoration – Rejecting the search for novelty
    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.
    Research output: Contribution to journal › Article › peer-review
  • PublishedReleasing grey squirrels into the wild
    Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390
    Research output: Contribution to journal › Article
  • PublishedRewilding needs to clarify the role of management of invasive species
    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438
    Research output: Contribution to journal › Letter
  • PublishedThe search for novelty continues for rewilding
    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585
    Research output: Contribution to journal › Article › peer-review
  • PublishedWhy the pine marten is not every red squirrel’s best friend
    Shuttleworth, C. & Hayward, M., 2019, The Conversation.
    Research output: Other contribution

2018

2017

2016

2015

2014

2013

2012

2010

2009

2008

2005

2004

2003

2002

2001

2000

1999

1997

1995

2022

  • Using remote activated cameras to estimate relative abundance and habitat preference of red squirrels

    Population estimates are essential for the conservation of wildlife. However, accurate estimates can be labour-intensive, particularly for small and medium-sized mammals where capture and handling of individuals is often required. Our study aimed to provide a robust and standardised method of determining red squirrel abundance using remotely activated cameras, while also quantifying how red squirrel abundance relates to key habitat characteristics. Fifty sample points were selected in four habitat categories in forests across Anglesey. We found a strong correlation (r = 0.78) between the number of red squirrels live-trapped over ten-days with the number of camera images recorded during a five-day period. Red squirrel abundance was negatively related to stand canopy openness, while the presence of Scots pine and increased tree species diversity were positively related to squirrel abundance. Our results demonstrate that remotely activated cameras can provide a suitable tool for surveying red squirrel abundance and guiding habitat management.

    19 Dec 2022

    Activity: Oral presentation (Speaker)

2021

2019

  • Win or bust - Returning the red squirrel to Snowdonia

    One woman's passion for the wilds of North Wales has ignited a resurgence of the charismatic red squirrel. Ecologist Dr Craig Shuttleworth tells us more about the return of red squirrels and their friends the pine martens to Snowdonia National Park.

    3 Jan 2019

    Links:

    Activity: Types of Public engagement and outreach - Media article or participation (Contributor)
  • Captive Red Squirrel Management: National Recommendations & Guidance

    Zoological red squirrel collections provide publicly exhibited specimens which are frequently the only opportunity for communities to see this rare native species and to learn about the threats posed to remnant wild populations. Collections are also an important resource for conservation translocation.

    This guidance builds upon historical recommendations for the management and husbandry of captive collections1, evolves the co-ordination of the national studbook and integrates key IUCN guidance2 on the selection of animals for any proposed wild release.

    The maintenance of an effective studbook requires agreement and mutual co-operation. Although this document presents a current best practice, we recognise that management must be adaptive and consequently that the text is a foundation for collaboration and the sharing of future lessons learned and knowledge.

    2019

    Activity: Types of Public engagement and outreach - Media article or participation (Contributor)

I am currently managing the Heritage Fund funded five year Magical Mammals applied conservation project in north Wales. This aims to widen participation and ldearning within mammal conservation and translocation programmes.